John Santiago and Brady Oliveira were the two most productive running backs in UND's Division I history, which dates back to 2008.

In Division I, the two were Nos. 1 and 2 in career rushing attempts, career rushing yards and Nos. 1 and 3 in career rushing touchdowns and career all-purpose yards.

Santiago and Oliveira -- both now with the Winnipeg Blue Bombers of the Canadian Football League -- dominated the Fighting Hawks' backfield for nearly all of Bubba Schweigert's tenure as head coach.

So it certainly feels like a new era at Memorial Stadium, where the Fighting Hawks went through the fifth practice of the 2019 fall camp Wednesday morning.

"I mean, it's tough to lose two All-Americans like that," UND running back James Johannesson said. "They did a good job teaching us how to lead the team. But we do miss having them here."

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The effort to replace Santiago and Oliveira starts with Johannesson, a 6-foot-1 transfer from the University of Minnesota.

In 2017, Johannesson had 41 carries for 248 yards and four touchdowns. He bumped that production to 93 carries for 651 yards and four touchdowns in 2018.

For his career, the former Fargo South star is averaging 6.7 yards per carry.

He was dominant in a three-game stretch during the middle of 2018. He ran 22 times for 141 yards at Northern Colorado on Sept. 29, then ran 16 times for 108 yards against Montana and had a breakout performance the following week at Sacramento State, where he ran 11 times for 213 yards.

He broke Dave Osborn's school record for yards per carry at Sacramento State (19.36).

One of the few knocks on Johannesson at UND has been his durability, an area he tried to address this offseason. He slimmed down from a bulky 250 pounds to 240.

"I feel like I have more stamina, and I'm not getting tired after the third or fourth play," Johannesson said. "I feel great. I've been rehabbing both my hamstrings. I'm doing a much better job this year taking care of myself than I have in the past."

Johannesson is listed No. 1 on the depth chart this fall, followed by sophomore speedster Cam McKinney and senior Austin Gordon.

Gordon's roller-coaster career has taken him from a walk-on showing promise at running back to getting suspended for an entire season by the NCAA to switching to linebacker and now moving back to playing offense.

The Maple Grove, Minn., native hasn't seen extensive action at running back since his redshirt freshman season. In 2016, Gordon had 33 carries for 243 yards. His 7.4 yards per carry led the team.

But suspension and injury derailed Gordon's playing time. Now, he's happy to be back counted on offensively.

"It feels more like home," Gordon said. "I had fun on the defensive side, but it was something I'm definitely not used to. It's nice to be back what I'm used to doing and feel like I'm best at doing."

Gordon said his injured ankle is nearly as strong as before the injury.

"At first, it was really slow trying to get everything back acclimated to moving and putting weight on it and what not," Gordon said. "The trainers did a good job getting the ankle strength close to where it was before the injury.

"I'm really excited for the season. Me and James have been close since he got here. We worked out together over the summer and came to the field just us sometimes. We have a good bond, and I'm excited to see what we can do with the new offense and the opportunities it can create for the both of us."